In Puritan New England, Hester Prynne (Lillian Gish) marries Roger Chillingsworth (Henry B. Walthall) at the behest of her father. When Roger is away, Hester forms a bond with the sensitive Rev. Arthur Dimmesdale (Lars Hanson). Later, when Hester gives birth to a daughter, people in the village suspect that Roger is not the father and force Hester to wear a letter A for adultery until she reveals the father of her child. Arthur, meanwhile, is overcome by guilt and decides to take action.
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